The Evolution of Vertebrate Design by Leonard B. Radinsky
Author:Leonard B. Radinsky [Radinsky, Leonard B.]
Language: eng
Format: epub
Publisher: The University of Chicago Press
Published: 1987-11-27T16:00:00+00:00
Fig. 10.1 Phylogeny of living and extinct amphibians.
The relatively ponderous skeleton and short limbs (fig. 10.2) suggest that the majority of the labyrinthodonts were slow, clumsy walkers on land, and they probably were truly amphibious and spent much of their time lying about in shallow water, like modern crocodiles. The long jaws and sharp conical teeth indicate that they were predators, and the large palate teeth suggest that they fed on relatively large prey—probably fishes and possibly on each other. Some of the smallest labyrinthodonts were relatively lightly built, proportioned like modern lizards, and, like lizards, they probably fed mainly on terrestrial invertebrates (insects and the like). During the first 100 million years of their history, some lines of labyrinthodonts became better adapted for land life. They developed stronger limbs and vertebral columns and probably were completely terrestrial predators. Only one group (the diadectids) appears to have become herbivorous. We make this inference from their large bodies (up to 10 feet in length), their very small heads, and small, blunt, transversely widened teeth. Flattened teeth are characteristic of living herbivores from elephants to fish. The shape functions to increase the surface area available for grinding down tough plant material. The length of the jaws is highly correlated with the length of the head. As we learned in chapter 8, the length of the jaw is the out-lever in the system producing out-force at the tooth row. Shortening the out-lever is one biomechanical way to increase the out-force. Large forces are needed to break down tough plant material. It is surprising that only one group of amphibians became herbivorous, since there was an abundance of vegetation on land.
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